The MIKE is the middle linebacker, who is generally more of a run-stopper than someone used in coverage. The MIKE is also responsible for calling out the plays to the defense and helps the defensive linemen get properly aligned. For the Buccaneers, Bruce Carter (No. 50) and Kwon Alexander (No. 58) have both seen time as the starting MIKE linebacker.
The WIL linebacker is the weak side outside linebacker. Lavonte David (No. 54) fills that role for the Bucs. The WIL is usually the fastest of the three linebackers and can cover a lot of ground. Coverage is also a primary responsibility of the WIL.