Tough question. Penn State is replacing All-American defensive end Carl Nassib, starting quarterback Christian Hackenberg and two other key starters -- Anthony Zettel and Jordan Lucas. But nose tackle Austin Johnson did some things that few players at his position could. And with the Lions inexperienced at the D-tackle positions, he gets my vote as the biggest personnel loss.
Penn State offensive coordinator Joe Moorhead talks to his players during spring practice on April 6, 2016 on the Lasch fields. Joe Hermitt, PennLive
Penn State's season concludes with the Blue-White game Saturday at Beaver Stadium.