Flat and listless, Penn State traveled to 0-4 UCLA Saturday and fell behind 27-7 by halftime.
They rallied furiously in the second half but lost 42-37 as the Bruins' Nico Iamaleava riddled them for five touchdowns, three passing and two on the ground. It marked the first time a winless team with a record of 0-4 or worse had beaten a Top Ten opponent in 40 years, an upset by a 24.5-point underdog that had fired its head coach, offensive coordinator and defensive coordinator.
Tight ends coach Rick Neuheisel, son of CBS broadcaster and former Bruin coach Rick Neuheisel, called plays.